Topic on Talk:Genes
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Metabolic efficiency is the price you pay for genes that are helpful in other ways - so if you get an advantage (say you move faster) you pay for that by having lower metabolic efficiency and having to eat more food. But if you take on a detriment (e.g. you are a ugly) you get paid for that by having higher metabolic efficiency and eating less food.
If this was confusing, it might be worth looking into rewriting that line.